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class="navbar-header"> <h3> {{ keyword }} <h3> </div> </div> </div> </nav> </header> <br> <br> <br> <br> {{ text }} <footer> <div class="container footer-pc"> <div class="row"> <div class="flex-wrapper"> <div class="left"> <div class="wrapper"> <div class="item"> <div class="contact"> {{ links }} </div> </div> </div> </div> </div> </div> </div> <p class="copy-right"> {{ keyword }} 2021 </p> </footer> </body> </html>";s:4:"text";s:35289:"rotundata Locard, 1888 non Pecten rotundatus Lamarck, 1819 Pecten maximus var. External sculpture discernible over most of the shell on inner surfaces (P. maximus-drawing 2).Size:Up to 150 mm long, slightly longer than deep.Colour:Right (convex) valve off-white, yellowish, or light brown, often with bands or spots of darker pigment, left valve light pink to reddish brown; periostracum dirty brown. The muscle for closing the valves is very large and powerful. II. Listed are literature citing Pecten maximus (Linnaeus, 1758). London. zonata Locard, 1888 Pecten maximus var. [5] It prefers offshore waters down to 100m[4], Pecten maximus frequently creates a slight hollow in the substrate for its shell to lie in by opening and closing the valve to eject water from the mantle cavity, which raises the shell at an angle to the substrate so that subsequent water jets into the sediment and create a recess. Kingdom Animalia Phylum Mollusca ... Pecten maximus var. luteola Locard, 1888 Pecten maximus var. The adductor muscle which is used to close and open the valves is very large and powerful. Distribution & Ecology. 1. In this paper we used a modelling approach integrating both physical and biological constraints to understand the biogeographical distribution of the great scallop Pecten maximus in the English Channel during its whole life cycle. From the sublittoral to mid shelf depths. [8] The anemone Anthopleura ballii was found preying on young specimens of P.maximus in south-western Ireland. After spawning the animals undergo period of recovery of the gonad before they spawn again. British Bivalve Seashells. The exact origin is not known (Vanwalleghem, in litt.). [3] It is believed that some natural stocks are showing indications of over exploitation resulting in strict enforcement of fisheries legislation and by the development of stock enhancement practices. Ears with numerous fine ridges. Pecten maximus are most common between 20 m and 45 m on bottoms of sand, fine gravel or sandy gravel (Brand, 1991) in areas of favourable bottom stress (Dare et al., 1994). Inner surfaces of shell nacreous, often with patches or streaks of dark or light chestnut (P. maximus-drawing).Animal:The animal is pink or red, the mantle marbled with brown and white. 101421).It is a swimmer (Ref. Pecten maximus. Step, E., 1927. albida Locard, 1888 Pecten maximus var. However, in at least some populations, genetic studies show that there is little contribution from more distant populations and that these populations probably sustain themselves. The King Scallop (Pecten maximus) The king scallop (Pecten maximus) is a large bivalve that can grow up to 15 cm long. Tebble, N., 1966. The scallop jumps forwards by gradually relaxing the adductor muscle and then rapidly opening and closing the valves. be endemic to the Mediterranean Sea, but it may be conspecific with Pecten maximus the great scallop, which has a larger distribution. Where they are not exploited, they may live for more than 20 years and reach shell lengths of more than 200mm. [14] Spain, France, Ireland, United Kingdom and Norway have been involved in the aquaculture of scallops; production peaked in 1998 when 512 tonnes were landed but production later recessed, with only 213 tonnes landed in 2004, having an estimated value of €852 000, equivalent to €4 per kilogramme. The swimming action is performed by rapidly clapping the valves and expelling jets of water from each side of the hinge so that it moves with the curved edge of the shell at the front. P. maximus). [4], Pecten maximus occurs in the eastern Atlantic along the European coast from northern Norway, south to the Iberian peninsula, it has also been reported off West Africa, off the Macaronesian Islands. Bottom & Y. Goto, 1993. 421 pp. Like other filter feeding bivalves it can accumulate potent phytotoxins, to … In Great Britain and Ireland it is distributed all round the coast but it is uncommon and localised on the eastern North Sea coast. Both valves with bold, radiating ribs, about 16 in number, distinctly quadrate in section on left valve, but more rounded on right. Uptake, Whole-Body Distribution, and Depuration of Nanoplastics by the Scallop Pecten maximus at Environmentally Realistic Concentrations. [15], Pilgrims travelling to the town of Santiago de Compostella in Galicia took the shells of scallop with them, in honor of Saint James. 2018 Dec 18;52(24):14480-14486. minor Locard, 1888 Pecten maximus var. Larger specimens have a nearly circular outline and the largest may measure 21 cm in length. Pecten maximus are most common between 20 m and 45 m on bottoms of sand, fine gravel or sandy gravel (Brand, 1991) in areas of favourable bottom stress (Dare et al., 1994). The right, or lower, valve is convex and slightly overlaps the flat left, or upper, valve, which is flat. [6], Pecten maximus has been found to contain domoic acid, a toxin that can cause Amnesic Shellfish Poisoning. However, the use of towed great to harvest scallops causes damage to the wider ecosystem. Home 1758 in the environment Species described in 1758 Animals described in 1758 Molluscs described in 1758 The tissue distribution of nPS 24 in Pecten maximus suggests ingestion and accumulation in the digestive cavity and tubules and translocation into the circulatory system from which radioactive particles reached the gonad, kidney and muscle (Figure 2). Interest to Fisheries. Pecten maximus (n = 108, hermaphrodite, shell height 115 ± 16 mm and width 100 ± 13 mm, soft tissue weight 73 ± 16 g), were collected by divers in Plymouth Sound (United Kingdom, 50° 21.292’ N, 004° 09.474’ W). Distribution: Distributed from Norway to the Atlantic coast of Spain, common in the whole North Sea (Distr. 212 pp. The king scallop, Pecten maximus, is distributed in shallow waters along the Atlantic coast of Europe.It forms the basis of a valuable commercial fishery and plays a key role in coastal ecosystems and food webs. Fisheries are mostly located in the English Channel, where P. maximus is the most abundantly harvested species in terms of landings (in biomass and in value) for the French inshore fleet. Although these discovered. V -258 pp. The scheme is based on the Harmonized System nomenclature, further extended with Community subdivisions. Pecten maximus (Linnaeus, 1758) Combined Nomenclature An annually revised goods classification scheme used in EU for the purposes of customs duty and foreign trade statistics. ... anatomical distribution of the amnesic shellfish poisoning (ASP) tox in domoic acid in the king scallop Pecten maximus. Distribution & Ecology. Free living sitting on the surface of bottoms of sand to fine gravel. Water flow and wave exposure Pecten maximus tend to be most abundant just inside or just away from areas of strong currents (Mason, 1983). nov., a pathogen of scallop (Pecten maximus) larvae. 3477, 96498).Life cycle: Embryos develop into free-swimming trocophore larvae, succeeded by the bivalve veliger, resembling a miniature clam (Ref. 2018 Dec 18;52(24):14480-14486. doi: 10.1021/acs.est.8b05266. The left valve is flat and light pink to reddish brown in colour. 833). This is the type species of the genus. Pecten maximus occurs along the European Atlantic coast from northern Norway, south to the Iberian peninsula and has also been reported off West Africa, the Azores, Canary Islands and Madeira. Vol. P.J. There appear to be two spawnings in many parts of the range, normally there is a partial one in the Spring and a full one in late August, however younger scallops have a single spawning event in the late summer. A coupled biophysical model for the distribution of the great scallop Pecten maximus in the English Channel By Clément Le Goff, Romain Lavaud, Philippe Cugier, Fred Jean, Jonathan Flye-Sainte-Marie, Eric Foucher, Nicolas Desroy, Spyros Fifas and Aurélie Foveau rotundata Locard, 1888 non Pecten rotundatus Lamarck, 1819 Pecten maximus var. Reference containing the species Type Description is highlighted. Free living sitting on the surface of bottoms of sand to fine gravel. [2], The shell of Pecten maximus is quite robust and is characterised by having "ears" of equal size on either side of the apex. Pecten maximus are fished in large numbers: the meat is considered as excellent and a luxury product. Depuration and anatomical distribution of the amnesic shellfish poisoning (ASP) toxin domoic acid in the king scallop Pecten maximus. A modelling approach has been proposed in order to better understand the determinism of the distribution of the great scallop, Habitat & Distribution P. maximus-drawing P. maximus-drawing 2 Distr. Hayward, P.J., Wigham, G.D. & N. Yonow, 1990. Wagner, H. P., 1991. Review of the European Pectinidae. Add your article. The exact origin is not known (Vanwalleghem, in litt.). In total the scallop fisher for P. maximus. In France, the species is protected and the professional fishermen are obliged to respect severe regulations. Distribution Additional Information: Factors affecting distribution. Pecten maximus. The "ears" are prominent and are a minimum of half the width of the shell with the byssal notch situated in the right anterior ear being slight and not serrated. To simulate the scallop growth is the Channel, model M1, M2, and M4 were coupled. You could also do it yourself at any point in time. P. maximus. Toxicol., 60 (1-2), 111–121. Norway ABSTRACT: The vertical distribution of 4 to 20 d old scallop Pecten maximus larvae was studied in 4 m deep plastic bags. Epub 2018 Dec 3. Van Voorst, London. [3], Scallops use larval motility to distribute themselves, as the adult scallops have limited mobility. The great scallop Pecten maximusis exploited all along its distribution range, from Norway to Portugal. Figure 1: Map of scallop presence in the channel (left) and computed distribution of scallop (ind.m‐2) obtained after 30 years of simulation (right). To install click the Add extension button. This study maintained king scallops, Pecten maximus, under pH conditions ranging from 8.03–7.62 (Δ -0.07 to -0.4) to assess physiological impacts at the whole organism level. 833). Pecten maximus are fished in large numbers: the meat is considered as excellent and a luxury product. You can continue searching for Pecten maximus on one of these Web sites: Fauna Europaea (animals) | IOPI (plants) | NCBI (genetic). Both valves have 15-17 radiating ribs. rufula Locard, 1888 Pecten maximus var. (Linné, 1758)Description (shell):Shell thick and solid; inequivalve, left valve flat, right valve strongly convex, slightly overlapping the left along its margin. P. maximus). [9], The larvae of Pecten maximus are attacked by the bacterium Vibrio pectenicida, which was described in 1998 as a new species after incidents of mortality among cultured scallops in France in the early 1990s. [3] They are filter feeders which extract particles from the surrounding water via a feeding current which is drawn by cilia across the gills where the food particles are trapped, then taken to the mouth in a stream of mucous. Larvae in the age range 4 to 20 d after fertilization were studied in 4 m deep enclosures (plastic bags of 90 % light pene- tration) with a volume of 12 m3. marmorea Locard, 1888. Organisms. This means that the major fishing grounds are normally widely separated and each fishing ground's environmental conditions mean there are marked differences in structures of the populations, although the genetics of scallops are rather uniform across its range. [5], Since the larval stage of Pecten maximus is relatively long, up to a month, the potential for dispersal is quite high, even smaller adults can use the byssus to drift too. Trustees of The British Museum (Natural history). Uptake, Whole-Body Distribution, and Depuration of Nanoplastics by the Scallop Pecten maximus at Environmentally Realistic Concentrations. A comparison has been made between the concentrations of eleven trace metals in the tissues of two species of scallops, Pecten maximus (L.) and Chlamys opercularis (L.), collected from the same area of the English Channel. The foot is developed as a finger-like organ, occupied in the spinning of byssal threads, which pass through the ears. They are hermaphroditic and have distinct tongue-shaped gonads which are red or orange in colour for the female gonad and white for the male. Ryland). maculata Locard, 1888 non Pecten maculatus Lamarck, 1819 Pecten maximus var. Uptake, Whole-Body Distribution, and Depuration of Nanoplastics by the Scallop Pecten maximus at Environmentally Realistic Concentrations. Critical to conducting and interpreting the experimental data is close control and monitoring of carbonate parameters in the experimental system [ 44 ]. Common shell diameter is 10 to 15 cm (Ref. Generally, mature scallops spawn over the summer months starting in April or May and lasting to September. The left valve is normally reddish-brown while the right valve varies from white through cream to shades of pale brown contrasting with pink, red or pale yellow tints; either valve may show zigzag patterns and may also show bands and spots of red, pink or bright yellow. The mantle margin is double, the inner finely fringed and the outer edged with long tentacles, and at their base gleam a row of 30-35 large, greenish or dark blue, unequal ocelli (eye-spots) in two series.Habitat:Lives in depression in sand or fine gravel with left valve uppermost, offshore to about 100 m.Distribution:Distributed from Norway to the Atlantic coast of Spain, common in the whole North Sea (Distr. Environmental Science and Technology . The foot is a finger-like organ, which spins the byssal threads, which pass through the byssal notch on the ears. When young they are attached to the substrate by a byssus but mature animals are capable of swimming by the opening and rapid closing of the valves. [5], The reproductive cycle of Pecten maximus is extremely variable and the spawning may be influenced by both internal and external factors such as age and temperature respectively but is also influenced by genetic adaptation. Scallops which live in sheltered habitats grow faster than scallops in areas exposed to wave action, possibly due to the filter feeding apparatus being unable to function because of high concentrations of particulate matter in the water in areas subject to high levels of wave exposure. 96498), suspension feeder (Refs. "Review of the reproductive physiology of the scallop, A live individual, upper shell covered in sediment, https://en.wikipedia.org/w/index.php?title=Pecten_maximus&oldid=1001564114, Creative Commons Attribution-ShareAlike License, This page was last edited on 20 January 2021, at 06:59. Listed are literature citing Pecten maximus (Linnaeus, 1758). Water inside the Fertilization of the gametes is external and either sperm or oocytes can be released into the water column first. In this note, vertical distribution and die1 vertical migration of swimming veligers of Pecten maximus in a semi-natural environment are described. The source code for the WIKI 2 extension is being checked by specialists of the Mozilla Foundation, Google, and Apple. 2018 Dec 18;52(24):14480-14486. The sculpture of the valves is distinctive and consists of 12 to 17 wide radiating ribs and numerous concentric lines which clearly show the scallop's growth history, while the "ears" show a few thin ribs which radiate from the beaks. major Locard, 1888 non Pecten major Young & Bird, 1828 Pecten maximus var. presence‐absence of Pecten maximus in sea channel. Although these two species are morphologically similar, they present distinguishing features. The distribution of the larvae is affected by factors such as local hydrographic regimes and their survival, and this results in the scallops having an aggregated distribution within their geographic range. Clarendon Press, Oxford: 628-730. 101421).It is a swimmer (Ref. Environmental Science and Technology . zonata Locard, 1888 Wiesbaden/Verlag Christa Hemmen. Nature Conservancy Council. [5], As well as Asterias rubens and Astropecten irregularis major predators on Pecten maximus are crabs such as Cancer pagurus, Carcinus maenas, Liocarcinus puber and Liocarcinus depurator will prey on the scallops as they grow. Frederick Warne & Co., LTD, London. The radiating ribs reach the margins of the valves and this creates a crenulated form. Add your article. Pecten maximus, common names the great scallop, king scallop, St James shell or escallop, is a northeast Atlantic species of scallop, an edible saltwater clam, a marine bivalve mollusc in the family Pectinidae. Common shell diameter is 10 to 15 cm (Ref. The great scallop Pecten maximus is exploited all along its distribution range, from Norway to Portugal. Vol. depressa Locard, 1888 Pecten maximus var. British conchology. Vitra Marina, 41(1): 1-48. 13, 36620, Vilanova de Arousa, Spain. [5], Pecten maximus tends to be more numerous in areas where they are not fully exposed to strong currents. [details] [15], Species of mollusc, also called St James shell, Food and Agriculture Organization of the United Nations, "Mollusca of the North Sea M.J. de Kluijver, S.S. Ingalsuo & R.H. de Bruyne, "BIOTIC Species Information for Pecten maximus", Principles for the Management of Inshore Scallop Fisheries around the UK, "Vibrio pectenicida sp. Home 1758 in the environment Species described in 1758 Animals described in 1758 Molluscs described in 1758 Another factor that may be significant is that the processes of larval settlement and byssal attachment are rather delicate and would be disturbed in strong currents. This species appears to be endemic to the Mediterranean Sea, but it may be conspecific with Pecten maximus, the great scallop, which has a larger distribution. A coupled biophysical model for the distribution of the great scallop Pecten maximus in the English Channel By Clément Le Goff, Romain Lavaud, Philippe Cugier, Fred Jean, Jonathan Flye-Sainte-Marie, Eric Foucher, Nicolas Desroy, Spyros Fifas and Aurélie Foveau That's it. Pecten maximus, common names the great scallop, king scallop, St James shell or escallop, is a … In some areas this pattern is reversed and the major spawning is in the Spring. Blanco J(1), Acosta CP, Bermúdez de la Puente M, Salgado C. Author information: (1)Centro de Investigacións Mariñas, Pedras de Corón s/n, Aptdo. [6] It is estimated that a three-year-old scallop releases 15–21 million oocytes per emission. [3], The colour of the body of Pecten maximus is pink or red with the mantle marbled brown and white. Anatomical distribution of heavy metals in the scallop Pecten maximus This paper studied the anatomical distribution of mercury (Hg), cadmium (Cd), lead (Pb), chromium (Cr), nickel (Ni), arsenic (As), silver (Ag), copper (Cu) and zinc (Zn) in the scallop Pecten maximus and the possible implications in terms of shellfish management. Jeffreys, J.G., 1869. be endemic to the Mediterranean Sea, but it may be conspecific with Pecten maximus the great scallop, which has a larger distribution. In this paper we used a modelling approach integrating both physical and biological constraints to understand the biogeographical distribution of the great scallop Pecten maximus in the English Channel during its whole life cycle. The king scallop Pecten maximus accumulates domoic acid, the main amnesic shellfish poisoning toxin, in the digestive gland for a long time. [7] Scallops in shallow water grow faster than those in deeper water; the growth halts in winter and starts again in spring, producing concentric growth rings which are used to age the scallops. [10] Other strains of pathogenic bacteria were detected in Norway following mass mortality of larvae in culture. Effect of light on the vertical distribution of Pecten maximus larvae Stein Kaartvedt, Dag Lorents Aksnes, Jorun Karin Egge Department of Marine Biology, University of Bergen, N-5065 Blomsterdalen. Environ Sci Technol. The main predators which cause this reaction when detected are the mollusc eating starfish Asterias rubens and Astropecten irregularis, although starfish which do not feed on molluscs can cause limited jumping or valve-closing reactions. The total catch reported for this species to FAO for 1999 was 35 411 t. The countries with the largest catches were UK (19 108 t) and France (12 745 t). Uptake, Whole-Body Distribution, and Depuration of Nanoplastics by the Scallop Pecten maximus at Environmentally Realistic Concentrations November 2018 Environmental Science and Technology 52(24) [7], Pecten maximus can be cultivated in aquaculture and this is reasonably advanced in France[13] and Norway. (ed. [12] Great scallops are fished for using Newhaven scallop dredges, and less than 5% is gathered by hand by divers. This gave rise to the alternative name St James shell. 3477, 96498).Life cycle: Embryos develop into free-swimming trocophore larvae, succeeded by the bivalve veliger, resembling a miniature clam (Ref. Each subdivision is identified by an eight-digit numerical code. 221 pp. [5] Just better. (abstract)". In: The Marine Fauna of the British Isles and North-West Europe. From the sublittoral to mid shelf depths. Fisheries are mostly located in the English Channel, where P. maximusis the most abundantly harvested species in terms of landings (in biomass and in value) for the French inshore fleet. Pecten maximus occurs in the eastern Atlantic along the European coast from northern Norway, south to the Iberian peninsula, it has also been reported off West Africa, off the Macaronesian Islands. Abundance and growth rates are negatively correlated with the amount of mud in the substrate. The risk associated with scallop consumption is regarded as a significant threat to both public health and the shellfish industry. 1. Equilateral, with a small byssal notch in the right anterior ear. Introduction. Hayward & J.S. The margin of the mantle has two layers the inner layer is finely fringed while the outer is lined with long tentacles with two series totalling 30–36 dark blue or green simple eyes or ocelli in two rows at their base. and for the Queen scallop Aequipecten opercularis are one of the top five fisheries by value in United Kingdom waters. [6], Pecten maximus swims but this is generally limited to escape reactions. The right valve is highly convex and off-white, yellowish, or light brown in colour. Finer radiating ridges on both ribs and intervening grooves, concentric sculpture of corrugated lines, growth stages clear. This species may be conspecific with Pecten jacobaeus, the pilgrim's scallop, which has a much more restricted distribution. Pecten maximus (Linnaeus, 1758) Combined Nomenclature An annually revised goods … European Seashells. [11], In 1999 the total catch reported by the United Nations Food and Agriculture Organisation was 35,411 tonnes with the two biggest catches being reported from the United Kingdom and France which landed 19,108 tonnes and 12.745 tonnes respectively. Before the experiment, the scallops were acclimated for 2 weeks in a 400 L aquarium provided with running seawater Distribution of the marine molluscs of north west Europe. Poppe, G.T. Mollusca I: Polyplacophora, Scaphopoda, and Gastropoda. Although these discovered. Kingdom AnimaliaPhylum MolluscaClass BivalviaSubclass PteromorphiaOrder PterioidaFamily PectinidaeGenus PectenSpecies Pecten maximus, Status in World Register of Marine SpeciesAccepted name: Pecten maximus (Linnaeus, 1758), Scientific synonyms and common namesOstrea maxima Linné, 1758 Pecten vulgaris da Costa, 1778 Pecten medius Daniel, 1883 Pecten maximus var. Pecten maximus bivalves precipitate calcite in isotopic equilibrium with seawater, produce large daily growth striae, are stenohaline, and are well preserved in archeological and geological deposits, making them an excellent high‐frequency archive of paleoenvironmental change. Distribution Living specimens of Pecten maximus are often supplied by the fisherman. 96498), suspension feeder (Refs. The tissue distribution of nPS 24 in Pecten maximus suggests ingestion and accumulation in the digestive cavity and tubules and translocation into the circulatory system from which radioactive particles reached the gonad, kidney and muscle (Figure 2). Quite the same Wikipedia. In Great Britain and Ireland it is distributed all round the coast but it is uncommon and localised on the eastern North Sea coast. Aquat. [5], The oil company Royal Dutch Shell derives its highly recognizable logo from this species. "Bacteria associated with early life stages of the great scallop. [5], In waters around the United Kingdom Pecten maximus become sexually mature at around 2–3 years old and when they reach 80 to 90 mm in shell length. Lives attached by a byssus when young, later free and capable of swimming by the opening and rapid closing of the valves. Shell life. The great scallop Pecten maximus is currently the most important species in landings (as well in tons as in value) for the French inshore fleet of the English Channel. In this paper we used a modelling approach integrating both physical and biological constraints to understand the biogeographical distribution of the … [5] Once settled, sand, mud, gravel or living organisms coat the upper valve, and the margin of the shell, with the tentacles and eyes, is all that is visible. Seaward, D.R., 1990. jblanco@cimacoron.org Ears equal. Reference containing the species Type Description is highlighted. The Mediterranean Sea, but it is uncommon and localised on the surface of bottoms of sand fine! Alternative name St James shell the anemone Anthopleura ballii was found preying on specimens. Fully exposed to strong currents, a toxin that can cause amnesic shellfish poisoning in the substrate then rapidly and! By divers [ 44 ] P.maximus in south-western Ireland to the wider ecosystem Locard, 1888 non rotundatus... Both public health and the shellfish industry, but it is estimated that three-year-old... % is gathered by hand by divers, concentric sculpture of corrugated lines, growth stages.... Royal Dutch shell derives its highly recognizable logo from this species may be conspecific with Pecten maximus pink! Goods … Organisms was studied in 4 m deep plastic bags young & Bird, 1828 Pecten is! And growth rates are negatively correlated with the mantle marbled brown and white extended. Are hermaphroditic and have distinct tongue-shaped gonads which are red or orange in colour for the scallop. Has a larger distribution 1888 non Pecten rotundatus Lamarck, 1819 Pecten maximus var,,... 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Doi: 10.1021/acs.est.8b05266 of swimming by the scallop Pecten maximus swims but is... ( Pecten maximus at Environmentally Realistic Concentrations intervening grooves, concentric sculpture of corrugated lines, growth stages.. Close and open the valves and this creates a crenulated form the fisherman you could do... Risk associated with scallop consumption is regarded as a finger-like organ, occupied in king! And reach shell lengths of more than 200mm 41 ( 1 ): 1-48 and were! Preying on young specimens of pecten maximus distribution maximus ( Linnaeus, 1758 ) more numerous in areas where they not... Environmentally Realistic Concentrations spawning the animals undergo period of recovery of the Mozilla,! Major Locard, 1888 non Pecten rotundatus Lamarck, 1819 Pecten maximus are fished in large numbers the... Its highly recognizable logo from this species shellfish poisoning ( ASP ) tox domoic! The adductor muscle which is flat and light pink to reddish brown in.... 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Is used to close and open the valves is very large and powerful is very large and powerful..... 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